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2. GWD-1-Q25 to Q28生理因素和环境因素对硅藻生物钟产生的影响
Recently biologists have been
interested in a tide-associated
periodic behavior displayed by
Line the diatom Hantzschia virgata, a
(5) microscopic golden-brown alga that
inhabits that portion of a shoreline
washed by tides (the intertidal zone).
Diatoms of this species, sometimes
called “commuter” diatoms, remain
(10) burrowed in the sand during high
tide, and emerge on the sand sur-
face during the daytime low tide.
Just before the sand is inundated by
the rising tide, the diatoms burrow
(15) again. Some scientists hypothesize
that commuter diatoms know that it
is low tide because they sense an
environmental change, such as an
alteration in temperature or a change
(20) in pressure caused by tidal move-
ment. However, when diatoms are
observed under constant conditions
in a laboratory, they still display
periodic behavior, continuing to bur-
(25) row on schedule for several weeks.
This indicates that commuter diatoms,
rather than relying on environmental
cues to keep time, possess an inter-
nal pacemaker or biological clock
(30) that enables them to anticipate peri-
odic changes in the environment.
A commuter diatom has an unusu-
ally accurate biological clock, a
consequence of the unrelenting
(35) environmental pressures to which
it is subjected; any diatoms that do
not burrow before the tide arrives
are washed away.提出现象及其研究
This is not to suggest that the
(40) period of this biological clock is
immutably fixed. Biologists have
concluded that even though a
diatom does not rely on the envi-
ronment to keep time, environmental
(45) factors—including changes in the
tide’s hydrostatic pressure, salin-
ity, mechanical agitation, and
temperature—can alter the period
of its biological clock according to
(50) changes in the tidal cycle. In short,
the relation between an organism’s
biological clock and its environment
is similar to that between a wristwatch
and its owner: the owner cannot
(55) make the watch run faster or slower,
but can reset the hands. However,
this relation is complicated in intertidal
dwellers such as commuter diatoms
by the fact that these organisms are
(60) exposed to the solar-day cycle as
well as to the tidal cycle, and some-
times display both solar-day and
tidal periods in a single behavior.
Commuter diatoms, for example,
(65) emerge only during those low tides
that occur during the day.环境因素对其生物钟的影响
现象解释
逻辑简图：
1P: recently biologist interested in T behavior of HV. Some hypothesis… however… indicate, rather than, possess..
2P: not suggest fix. Biologist conclude environmental factors can alter
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